Marine life  /  Blackwater & plankton  /  Heteropod
Reviewed 2026-09-05
Species entry

Heteropod

Carinariidae

A sea snail that gave up the bottom, turned its foot into a swimming fin, and became glass.

Size
To 50 cm as an adult; larvae a few millimetres
Depth
0–25 m
Habitat
Blackwater, open water at night
Active
Night, in open water
Range
Open ocean worldwide
Rarity
Lembeh Strait
3 stars of 5, a good week should turn one up
Heteropod veliger in black water, an orange body with long blue arms radiating from it, each arm lined with yellow ciliated bands.
Simon Buxton

Heteropods are predatory sea snails that live their whole lives in open water. The foot, which in a bottom-living snail is the muscular sole it crawls on, has become a single vertical fin that the animal beats while swimming upside down. The body is almost entirely transparent, so what is visible in a photograph is usually the gut, the eyes and the fin, with the rest inferred.

The Carinariidae keep a shell, but only a small glassy cap over the visceral mass, far too small to withdraw into and clearly on its way out — the related Pterotracheidae have lost it altogether. Carinaria cristata reaches half a metre and is one of the largest predatory gastropods in the ocean.

Most blackwater encounters are with larvae and juveniles of a few millimetres, at the veliger stage, when the animal still carries the coiled larval shell and the ciliated lobes it swims and feeds with. Larvae are not identifiable to species; the family is as far as a photograph will honestly take you.

Where to find:

LocationRarity Best monthsDepthHabitat
Lembeh Strait Indonesia editorial estimate
3 stars of 5, a good week should turn one up
Year-round
0–25 m
Blackwater, open water at night
Anilao Philippines editorial estimate
3 stars of 5, a good week should turn one up
Oct Nov Dec Jan Feb Mar Apr May
0–25 m
Blackwater, open water at night

Ratings are per location: 1 means found on most dives that look for it, 5 means a handful of records. Where enough dives have been logged the figure is calculated from them; otherwise it is an editorial estimate, and says so. We publish habitat and destination, not the dive site.

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